Cocoseae


Cocoseae is a tribe of cocosoid palms of the family Arecaceae.

Description

The fruit of the Cocoseae is a modified drupe, with a sclerenchymatous epicarp and a highly developed mesocarp, formed mainly by parenchyma. The endocarp is generally sclerenchymatous and protects the seeds from predation and drying. The most obvious synapomorphy of the species of this tribe is the presence, in the endocarp, of three or more "eyes" or pores of germination. According to Thomas C. Whitmore all Cocosoid palms have inflorescences in which the flowers are in triads.

Distribution

The Cocoseae are distributed mainly in the Neotropical regions, with two genera endemic to Africa and Madagascar, respectively.

Phylogeny

A 2016 molecular phylogenomic analysis of the tribe Cocoseae gives the following phylogenetic tree:

Systematics

The Cocoseae in the sense of Dransfield et al. are identified as natural relatives in most studies. Their systematic position within the Arecoideae is still unclear. Different studies see them as a sister group of the Reinhardtieae, as a sister group of the group from Reinhardtieae and Roystoneeae, or as a sister group of the Oranieae.
The tribe is divided into three subtribes. All three are monophyletic. Bactridinae and Elaeidinae are sister groups

Genera

SubtribeImageGenusExtant species
Beccariophoenicinae J.Dransf. & UhlBeccariophoenix Jum. & H.PerrierBeccariophoenix alfredii Rakotoarin., Ranariv. & J.Dransf. Beccariophoenix madagascariensis Jum. & H.Perrier Beccariophoenix fenestralis J. Dransfield & M. Rakotoarinivo
Butiinae SaakovAllagoptera NeesAllagoptera arenaria Kuntze Allagoptera brevicalyx M.Moraes Allagoptera campestris Kuntze Allagoptera caudescens Kuntze Allagoptera leucocalyx Kuntze
Butiinae SaakovButia Becc Butia archeri GlassmanButia arenicola BurretButia campicola NoblickButia capitata Becc.Butia catarinensis Noblick & LorenziButia eriospatha Becc. Butia exilata Deble & MarchioriButia exospadix NoblickButia lallemantii Deble & MarchioriButia lepidotispatha NoblickButia leptospatha NoblickButia marmorii NoblickButia matogrossensis Noblick & LorenziButia microspadix BurretButia odorata Noblick Butia paraguayensis L.H.Bailey Butia pubispatha Noblick & LorenziButia purpurascens GlassmanButia witeckii K.Soares & S. LonghiButia yatay Becc.
Butiinae SaakovJubaea KunthJubaea chilensis Baill.
Butiinae SaakovJubaeopsis Becc.Jubaeopsis afra Becc.
Butiinae SaakovCocos L.Cocos nucifera L.
Butiinae SaakovSyagrus Mart.
Butiinae SaakovParajubaea BurretParajubaea cocoides BurretParajubaea sunkha M.MoraesParajubaea torallyi Burret
Butiinae SaakovVoanioala J. Dransf.Voanioala gerardii J. Dransf.
Attaleinae DrudeAttalea Kunth
Elaeidinae Hook.f.Barcella Trail ex DrudeBarcella odora Drude
Elaeidinae Hook.f.Elaeis Jacq.Elaeis guineensis Jacq. Elaeis oleifera Cortés
Bactridinae DrudeAcrocomia Mart.Acrocomia aculeata Lodd. ex R.KeithAcrocomia crispa C. Baker ex. Becc.Acrocomia emensis LorenziAcrocomia glaucescens LorenziAcrocomia hassleri W.J.Hahn DrudeAcrocomia media O.F.CookAcrocomia totai Mart.
Bactridinae DrudeAiphanes Willd.
Bactridinae DrudeAstrocaryum G.Mey.
Bactridinae DrudeBactris Jacq. ex Scop.
Bactridinae DrudeDesmoncus Trail ex Drude
Bactridinae Drude---