Dryopidae


Dryopidae is a family of beetles, commonly named long-toed water beetles, in the superfamily Byrrhoidea. It was described by Gustaf Johan Billberg in 1820.

Description

Long-toed water beetles are named for their extended claws. Adults have dense hairs, which allow the beetles to breathe while underwater. The flight muscles of the females weaken as they age.
When the pupae complete the imago stage of their life cycle, they move towards running water, and may be attracted to lights.
Despite being referred to as aquatic insects, the beetles are unable to swim, clinging to detritus that float. All long-toed water beetles feed on plants that are in the water, but the larvae generally are terrestrial, and at least some also feed on plant matter. Stygoparnus is the only genus in the family in which both the larvae and adults are aquatic.
Genera of the family closely resemble riffle beetles, but the antennae are different from the long-toed water beetles, looking similar to clubs.

Habitat

Members of this family are found on every continent, except Antarctica and Australia, being more common in the tropics. In 2005, a possible fossilized specimen of Dryopidae was found in the Crato Formation by entomologists David Grimaldi and Michael S. Engel.
Beetles that have water as a habitat, including long-toed water beetles, can help show the quality of fresh water.

Genera

Ahaggaria Bollow, 1938Ceradryops Hinton, 1937Drylichus Heller, 1916Dryops Olivier, 1791Elmomorphus Sharp, 1888Elmoparnus Sharp, 1882Geoparnus Besuchet, 1978Guaranius Spangler, 1991Helichus Erichson, 1847Holcodryops Spangler, 1987Malaiseianus Bollow, 1940Onopelmus Spangler, 1980Oreoparvus Delève, 1965Pelonomus Erichson, 1847Phallodryops Delève, 1963Postelichus Nelson, 1989Protoparnus Sharp, 1883Pseudopelonomus Brown, 1981Quadryops Perkins & Spangler, 1985Rapnus Grouvelle, 1899Sostea Pascoe, 1860Sosteamorphus Hinton, 1936Strina Redtenbacher, 1867Stygoparnus Barr & Spangler, 1992Uenodryops Satô, 1981Palaeoriohelmis Bollow, 1940Potamophilites Haupt, 1956