Endomychidae


Endomychidae, or handsome fungus beetles, is a family of beetles with representatives found in all biogeographic realms. There are around 120 genera and 1300 species. The family was established based on the type genus Endomychus, a genus erected in 1795 by Panzer which was applied to a species that Linnaeus called Chrysomela coccinea. As the common name suggests, Endomychidae feed on fungi. Crowson, in his influential treatment of the beetles, placed the family within the Cucujoidea. They have a tarsal formal of 4-4-4 or 3-3-3 and the wings lack a closed radial cell. The second antennal segment has a sensory appendage that is as long as the third antennal segment. The family has also been grouped with the Coccinellidae in a group called the Trimera for having pseudotrimerous tarsi. A 2015 molecular phylogeny study found that the Cucujoidea were found to be non-monophyletic and the Endomychidae was refined with the removal of the Anamorphinae from within the family and elevated to the status of a full family, Anamorphidae. Mycetaeinae and Eupsilobiinae were also found not to belong within the clades of the core Endomychidae, and likewise reclassified into the families Mycetaeidae and Eupsilobiidae.
The subfamilies that are included:

Genera

These 70 genera belong to the family Endomychidae as defined in the past :Acinaces Gerstaecker, 1858 gAclemmysa Reitter, 1904 gAgaricophilus Motschulsky, 1838 gAmphisternus Germar, 1843 gAmphistethus Strohecker, 1964 gAncylopus Costa, 1854 gAnidrytus Gerstaecker, 1858 i c gAphorista Gorham, 1873 i c g bArchipines Strohecker, 1953 i c gAsymbius Gorham, 1896 gAtopomychus Tomaszewska & Szawaryn, 2013 gAtrichonota Arrow, 1925 gAvencymon Strohecker, 1971 gBeccariola Arrow, 1943 gBolbomorphus Gorham, 1888 gBrachytrycherus Arrow, 1920 gBystodes Strohecker, 1953 gCholovocera Motschulsky, 1838 gChondria Gorham, 1888 gCorynomalus Chevrolat in Dejean, 1836 i c gCyclotoma Mulsant, 1851 i gDanae Reiche, 1847 i c g bDanascelis Tomaszewska, 1999 i c gDapsa Latreille, 1829 gDexialia Sasaji, 1970 gDialexia Gorham, 1891 gDiscolomopsis Shockley, 2006 gDisplotera Reitter, 1877 gEctomychus Gorham, 1888 gEidorus gEndomychus Panzer, 1795 i c g bEphebus Chevrolat in Dejean, 1836 i c gEpipocus Germar, 1843 i c g bEpopterus Chevrolat in Dejean, 1836 i c gEucteanus Gerstaecker, 1857 gEucymon gEumorphus Weber, 1801 gGeoendomychus Lea, 1922 gGlesirhanis Shockley & Alekseev, 2014 gHadromychus Bousquet & Leschen, 2002 i c g bHoloparamecus Curtis, 1833 i c g bHylaia Guerin, 1857 gHylaperdina Tomaszewska, 2012 gIdiophyes Blackburn, 1896 gIndalmus Gerstaecker, 1858 gLeiestes Chevrolat, 1836 gLycoperdina Latreille, 1807 i c g bMeilichius Gerstaecker, 1857 gMerophysia Lucas, 1852 gMicroxenus Wallaston, 1861 gMycetina Mulsant, 1846 i c g bMychothenus Strohecker, 1953 gNatalinus Tomaszewska, 2011 gOhtaius Chûjô, 1938 gPalaeoestes Kirejtshuk & Nel, 2009 gPaniegena Heller, 1916 gParasymbius Arrow, 1920 gPhymaphora Newman, 1838 i c g bPhymaphoroides Motschoulsky, 1857 gPleganophorus Hampe, 1855 gPolymus Mulsant, 1846 gPseudindalmus Arrow, 1920 gRhanidea Strohecker, 1953 i c g bSaula Gerstaecker, 1858 i c gSinocymbachus Strohecker & Chujo, 1970 gSinopanamomus Esser, 2019 Stenotarsus Perty, 1832 i c g bStethorhanis Blaisdell, 1931 i c gTrochoideus Westwood, 1833 i c g bTrycherus Gerstaecker, 1857 gXenomycetes Horn, 1880 i c g b
Data sources: i = ITIS, c = Catalogue of Life, g = GBIF, b = Bugguide.net

Fossil genera

Burmalestes Tomaszewska and Ślipiński, 2018 Burmese amber, Myanmar, Late Cretaceous Cretolestes Tomaszewska, Ślipiński and Ren, 2018 Burmese amber, Myanmar, CenomanianCretaparamecus Tomaszewska, Ślipiński, Bai and Zhang, 2018 Burmese amber, Myanmar, CenomanianPalaeomycetes Tomaszewska, Ślipiński and Ren, 2018 Burmese amber, Myanmar, CenomanianZemyna Tomaszewska, 2018 '': Baltic amber, Eocene